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No, effects are not permanent. You can toggle them on or off as you please. In the campaign lobby screen you can press X to bring up a screen to see each skull you have collected and it will give you option enable or disable them.
Nope. The halo skulls actually have a meaning in Halo 3. They r for achievments and for score modifyers. When u find a skull, drop it quit (or continue) the level, then press X before u start a new game to see ur skull collection. Make sure u turn on campaign scoring to get more achivements and activating GOLD skulls gives u multiplyers for ur score (x2, x3, x4)